LOS ANGELES (AP) - Jerry Lewis, the manic, rubber-faced showman who jumped and hollered to fame in a lucrative partnership with Dean Martin, settled down to become a self-conscious screen auteur and found an even greater following as the tireless, teary host of the annual muscular dystrophy telethons, has died. He was 91.

Publicist Candi Cazau says Lewis died Sunday of natural causes in Las Vegas with his family by his side.

Tributes from friends, co-stars and disciples poured in immediately.

"That fool was no dummy. Jerry Lewis was an undeniable genius an unfathomable blessing, comedy's absolute!" Jim Carrey wrote Sunday on Twitter. "I am because he was!"

"The world has lost a true innovator & icon," comedian Dane Cook wrote.

In Las Vegas, a message honoring the comedian is being featured on a marquee at Caesars Palace, where Lewis was once a headliner and had also hosted telethons. In Los Angeles fans and admirers gathered at Lewis' two Hollywood Walk of Fame stars - one for television and one for film.

Martin & Lewis
Dean Martin and Jerry Lewis / Getty Images
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Lewis' career spanned the history of show business in the 20th century, beginning in his parents' vaudeville act at the age of 5. He was just 20 when his pairing with Martin made them international stars. He went on to make such favorites as "The Bellboy" and "The Nutty Professor," was featured in Martin Scorsese's "The King of Comedy" and appeared as himself in Billy Crystal's "Mr. Saturday Night."

"Jerry was a pioneer in comedy and film. And he was a friend. I was fortunate to have seen him a few times over the past couple of years. Even at 91, he didn't miss a beat. Or a punchline," Lewis' "The King of Comedy" co-star Robert De Niro said in a statement.

In the 1990s, he scored a stage comeback as the devil in the Broadway revival of "Damn Yankees." And after a 20-year break from making movies, Lewis returned as the star of the independent drama "Max Rose," released in 2016.

In his 80s, he was still traveling the world, working on a stage version of "The Nutty Professor." He was so active he would sometimes forget the basics, like eating, his associates would recall. In 2012, Lewis missed an awards ceremony thrown by his beloved Friars Club because his blood sugar dropped from lack of food and he had to spend the night in the hospital.

A major influence on Carrey and other slapstick performers, Lewis also was known as the ringmaster of the Labor Day Muscular Dystrophy Association, joking and reminiscing and introducing guests, sharing stories about ailing kids and concluding with his personal anthem, the ballad "You'll Never Walk Alone." From the 1960s onward, the telethons raised some $1.5 billion, including more than $60 million in 2009. He announced in 2011 that he would step down as host, but would remain chairman of the association he joined some 60 years ago.

"Though we will miss him beyond measure, we suspect that somewhere in heaven, he's already urging the angels to give 'just one dollar more for my kids,'" said MDA Chairman of the Board R. Rodney Howell on Sunday.

His fundraising efforts won him the Jean Hersholt Humanitarian Award at the 2009 Oscar telecast. But the telethon was also criticized for being mawkish and exploitative of children, known as "Jerry's Kids." A 1960s muscular dystrophy poster boy, Mike Ervin, later made a documentary called "The Kids Are All Alright," in which he alleged that Lewis and the Muscular Dystrophy Association had treated him and others as objects of pity rather than real people.

"He and his telethon symbolize an antiquated and destructive 1950s charity mentality," Ervin wrote in 2009.

Responded Lewis: "You don't want to be pitied because you're a cripple in a wheelchair, stay in your house!"

Lewis also sassed and snarled at critics and interviewers who displeased him. He pontificated on talk shows, lectured to college students and compiled his thoughts in the 1971 book "The Total Film-Maker."

"I am not ashamed or embarrassed at how seemingly trite or saccharine something in my films will sound," he wrote. "I really do make films for my great-great-grandchildren and not for my fellows at the Screen Directors Guild or for the critics."

In his early movies, Lewis played loose-limbed, buck-toothed, overgrown adolescents, trouble-prone and inclined to wail when beset by enemies. American critics recognized the comedian's popular appeal but not his aspirations to higher art; the French did. Writing in Paris' Le Monde newspaper, Jacques Siclier praised Lewis' "apish allure, his conduct of a child, his grimaces, his contortions, his maladjustment to the world, his morbid fear of women, his way of disturbing order everywhere he appeared."

TCM Classic Film Festival - Jerry Lewis Hand And Footprint Ceremony
Getty Images
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The French government awarded Lewis the Chevalier of the Legion of Honor in 1983 and Commander of Arts and Letters the following year.

Lewis had teamed up with Martin after World War II, and their radio and stage antics delighted audiences, although not immediately. Their debut, in 1946 at Atlantic City's 500 Club, was a bust. Warned by owner "Skinny" D'Amato that they might be fired, Martin and Lewis tossed the script and improvised their way into history. New York columnists Walter Winchell and Ed Sullivan came to the club and raved over the sexy singer and the berserk clown.

Lewis described their fledgling act in his 1982 autobiography, "Jerry Lewis in Person": "We juggle and drop a few dishes and try a few handstands. I conduct the three-piece band with one of my shoes, burn their music, jump offstage, run around the tables, sit down with the customers and spill things while Dean keeps singing."

Hollywood producer Hal Wallis saw them at New York's Copacabana and signed them to a film contract. Martin and Lewis first appeared in supporting roles in "My Friend Irma" and "My Friend Irma Goes West." Then they began a hit series of starring vehicles, including "At War With the Army," ''That's My Boy" and "Artists and Models."

But in the mid-1950s, their partnership began to wear. Lewis longed for more than laughs. Martin had tired of playing straight man and of Lewis' attempts to add Chaplinesque pathos. He also wearied of the pace of films, television, nightclub and theater appearances, benefits and publicity junkets on which Lewis thrived. The rift became increasingly public as the two camps sparred verbally.

"I knew we were in trouble the day someone gave Jerry a book about Charlie Chaplin," Martin cracked.

On July 24, 1956, Martin and Lewis closed shop, at the Copa, and remained estranged for years. Martin, who died in 1995, did make a dramatic, surprise appearance on Lewis' telethon in 1976 (a reunion brokered by mutual pal Frank Sinatra), and director Peter Bogdonavich nearly persuaded them to appear in a film together as former colleagues who no longer speak to each other. After Martin's death, Lewis said the two had again become friendly during his former partner's final years and he would repeatedly express his admiration for Martin above all others.

The entertainment trade at first considered Martin the casualty of the split, since his talents, except as a singer, were unexplored. He fooled his detractors by cultivating a comic, drunken persona, becoming star of a long-running TV variety show and a respected actor in such films as "Some Came Running," ''The Young Lions" and "Rio Bravo."

Lewis also distinguished himself after the break, revealing a serious side as unexpected as Martin's gift for comedy.

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